The Evolution of Combinatorial Phonology Willem Zuidema, Bart de Boer Abstract: A fundamental, universal property of human language is that its phonology is combinatorial. That is, one can identify a set of basic, distinct units (phonemes, syllables) that can be productively combined in many di-b®erent ways. In this paper, we review a number of theories-A and models that have been developed to explain the evolutionary transition from holistic to combinatorial signal systems, but -b¯nd that-A in all problematic linguistic assumptions are made, or crucial components of evolutionary explanations are omitted. We present a novel model to investigate the hypothesis that combinatorial phonology results from optimising signal systems for perceptual distinctiveness. Our model di-b®ers from previous models in two-A important respects. First, signals are modelled as trajectories through acoustic space. Hence, both holistic and combinatorial signals have a temporal structure. Second, we use the methodology from evolutionary game theory. Crucially, we show a path of ever increasing -b¯tness from holistic to combinatorial signals, where every innovation-A represents an advantage even if no-one else in a population has yet obtained it. Keywords: evolution of language, evolutionary game theory, combinatorial phonology